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<p>I was delayed here five days, and employed myself in examining the geology of the surrounding country, which was very interesting. We here see at the bottom of the cliffs, beds containing sharks teeth and seashells of extinct species, passing above into an indurated marl, and from that into the red clayey earth of the Pampas, with its calcareous concretions and the bones of terrestrial quadrupeds. This vertical section clearly tells us of a large bay of pure saltwater, gradually encroached on, and at last converted into the bed of a muddy estuary, into which floating carcasses were swept. At Punta Gorda, in Banda Oriental, I found an alternation of the Pampaean estuary deposit, with a limestone containing some of the same extinct seashells; and this shows either a change in the former currents, or more probably an oscillation of level in the bottom of the ancient estuary. Until lately, my reasons for considering the Pampaean formation to be an estuary deposit were, its general appearance, its position at the mouth of the existing great river the Plata, and the presence of so many bones of terrestrial quadrupeds: but now Professor Ehrenberg has had the kindness to examine for me a little of the red earth, taken from low down in the deposit, close to the skeletons of the mastodon, and he finds in it many infusoria, partly saltwater and partly freshwater forms, with the latter rather preponderating; and therefore, as he remarks, the water must have been brackish. <abbr>M.</abbr> <abbr epub:type="z3998:given-name">A.</abbr> dOrbigny found on the banks of the Parana, at the height of a hundred feet, great beds of an estuary shell, now living a hundred miles lower down nearer the sea; and I found similar shells at a less height on the banks of the Uruguay; this shows that just before the Pampas was slowly elevated into dry land, the water covering it was brackish. Below Buenos Aires there are upraised beds of seashells of existing species, which also proves that the period of elevation of the Pampas was within the recent period.</p> <p>I was delayed here five days, and employed myself in examining the geology of the surrounding country, which was very interesting. We here see at the bottom of the cliffs, beds containing sharks teeth and seashells of extinct species, passing above into an indurated marl, and from that into the red clayey earth of the Pampas, with its calcareous concretions and the bones of terrestrial quadrupeds. This vertical section clearly tells us of a large bay of pure saltwater, gradually encroached on, and at last converted into the bed of a muddy estuary, into which floating carcasses were swept. At Punta Gorda, in Banda Oriental, I found an alternation of the Pampaean estuary deposit, with a limestone containing some of the same extinct seashells; and this shows either a change in the former currents, or more probably an oscillation of level in the bottom of the ancient estuary. Until lately, my reasons for considering the Pampaean formation to be an estuary deposit were, its general appearance, its position at the mouth of the existing great river the Plata, and the presence of so many bones of terrestrial quadrupeds: but now Professor Ehrenberg has had the kindness to examine for me a little of the red earth, taken from low down in the deposit, close to the skeletons of the mastodon, and he finds in it many infusoria, partly saltwater and partly freshwater forms, with the latter rather preponderating; and therefore, as he remarks, the water must have been brackish. <abbr>M.</abbr> <abbr epub:type="z3998:given-name">A.</abbr> dOrbigny found on the banks of the Parana, at the height of a hundred feet, great beds of an estuary shell, now living a hundred miles lower down nearer the sea; and I found similar shells at a less height on the banks of the Uruguay; this shows that just before the Pampas was slowly elevated into dry land, the water covering it was brackish. Below Buenos Aires there are upraised beds of seashells of existing species, which also proves that the period of elevation of the Pampas was within the recent period.</p>
<p>In the Pampaean deposit at the Bajada I found the osseous armour of a gigantic armadillo-like animal, the inside of which, when the earth was removed, was like a great cauldron; I found also teeth of the Toxodon and Mastodon, and one tooth of a Horse, in the same stained and decayed state. This latter tooth greatly interested me,<a href="endnotes.xhtml#note-74" id="noteref-74" epub:type="noteref">74</a> and I took scrupulous care in ascertaining that it had been embedded contemporaneously with the other remains; for I was not then aware that amongst the fossils from Bahia Blanca there was a horses tooth hidden in the matrix: nor was it then known with certainty that the remains of horses are common in North America. <abbr epub:type="z3998:name-title">Mr.</abbr> Lyell has lately brought from the United States a tooth of a horse; and it is an interesting fact, that Professor Owen could find in no species, either fossil or recent, a slight but peculiar curvature characterizing it, until he thought of comparing it with my specimen found here: he has named this American horse <i epub:type="z3998:taxonomy">Equus curvidens</i>. Certainly it is a marvellous fact in the history of the Mammalia, that in South America a native horse should have lived and disappeared, to be succeeded in after-ages by the countless herds descended from the few introduced with the Spanish colonists!</p> <p>In the Pampaean deposit at the Bajada I found the osseous armour of a gigantic armadillo-like animal, the inside of which, when the earth was removed, was like a great cauldron; I found also teeth of the Toxodon and Mastodon, and one tooth of a Horse, in the same stained and decayed state. This latter tooth greatly interested me,<a href="endnotes.xhtml#note-74" id="noteref-74" epub:type="noteref">74</a> and I took scrupulous care in ascertaining that it had been embedded contemporaneously with the other remains; for I was not then aware that amongst the fossils from Bahia Blanca there was a horses tooth hidden in the matrix: nor was it then known with certainty that the remains of horses are common in North America. <abbr epub:type="z3998:name-title">Mr.</abbr> Lyell has lately brought from the United States a tooth of a horse; and it is an interesting fact, that Professor Owen could find in no species, either fossil or recent, a slight but peculiar curvature characterizing it, until he thought of comparing it with my specimen found here: he has named this American horse <i epub:type="z3998:taxonomy">Equus curvidens</i>. Certainly it is a marvellous fact in the history of the Mammalia, that in South America a native horse should have lived and disappeared, to be succeeded in after-ages by the countless herds descended from the few introduced with the Spanish colonists!</p>
<p>The existence in South America of a fossil horse, of the mastodon, possibly of an elephant,<a href="endnotes.xhtml#note-75" id="noteref-75" epub:type="noteref">75</a> and of a hollow-horned ruminant, discovered by <abbr epub:type="z3998:name-title">MM.</abbr> Lund and Clausen in the caves of Brazil, are highly interesting facts with respect to the geographical distribution of animals. At the present time, if we divide America, not by the Isthmus of Panama, but by the southern part of Mexico<a href="endnotes.xhtml#note-76" id="noteref-76" epub:type="noteref">76</a> in <abbr>lat.</abbr> 20°, where the great tableland presents an obstacle to the migration of species, by affecting the climate, and by forming, with the exception of some valleys and of a fringe of low land on the coast, a broad barrier; we shall then have the two zoological provinces of North and South America strongly contrasted with each other. Some few species alone have passed the barrier, and may be considered as wanderers from the south, such as the puma, opossum, kinkajou, and peccari. South America is characterized by possessing many peculiar gnawers, a family of monkeys, the llama, peccari, tapir, opossums, and, especially, several genera of Edentata, the order which includes the sloths, anteaters, and armadilloes. North America, on the other hand, is characterized (putting on one side a few wandering species) by numerous peculiar gnawers, and by four genera (the ox, sheep, goat, and antelope) of hollow-horned ruminants, of which great division South America is not known to possess a single species. Formerly, but within the period when most of the now existing shells were living, North America possessed, besides hollow-horned ruminants, the elephant, mastodon, horse, and three genera of Edentata, namely, the Megatherium, Megalonyx, and Mylodon. Within nearly this same period (as proved by the shells at Bahia Blanca) South America possessed, as we have just seen, a mastodon, horse, hollow-horned ruminant, and the same three genera (as well as several others) of the Edentata. Hence it is evident that North and South America, in having within a late geological period these several genera in common, were much more closely related in the character of their terrestrial inhabitants than they now are. The more I reflect on this case, the more interesting it appears: I know of no other instance where we can almost mark the period and manner of the splitting up of one great region into two well-characterized zoological provinces. The geologist, who is fully impressed with the vast oscillations of level which have affected the earths crust within late periods, will not fear to speculate on the recent elevation of the Mexican platform, or, more probably, on the recent submergence of land in the West Indian Archipelago, as the cause of the present zoological separation of North and South America. The South American character of the West Indian mammals<a href="endnotes.xhtml#note-77" id="noteref-77" epub:type="noteref">77</a> seems to indicate that this archipelago was formerly united to the southern continent, and that it has subsequently been an area of subsidence.</p> <p>The existence in South America of a fossil horse, of the mastodon, possibly of an elephant,<a href="endnotes.xhtml#note-75" id="noteref-75" epub:type="noteref">75</a> and of a hollow-horned ruminant, discovered by <abbr epub:type="z3998:name-title">MM.</abbr> Lund and Clausen in the caves of Brazil, are highly interesting facts with respect to the geographical distribution of animals. At the present time, if we divide America, not by the Isthmus of Panama, but by the southern part of Mexico<a href="endnotes.xhtml#note-76" id="noteref-76" epub:type="noteref">76</a> in <abbr>lat.</abbr> 20°, where the great tableland presents an obstacle to the migration of species, by affecting the climate, and by forming, with the exception of some valleys and of a fringe of low land on the coast, a broad barrier; we shall then have the two zoological provinces of North and South America strongly contrasted with each other. Some few species alone have passed the barrier, and may be considered as wanderers from the south, such as the puma, opossum, kinkajou, and peccari. South America is characterized by possessing many peculiar gnawers, a family of monkeys, the llama, peccari, tapir, opossums, and, especially, several genera of Edentata, the order which includes the sloths, anteaters, and armadilloes. North America, on the other hand, is characterized (putting on one side a few wandering species) by numerous peculiar gnawers, and by four genera (the ox, sheep, goat, and antelope) of hollow-horned ruminants, of which great division South America is not known to possess a single species. Formerly, but within the period when most of the now existing shells were living, North America possessed, besides hollow-horned ruminants, the elephant, mastodon, horse, and three genera of Edentata, namely, the Megatherium, Megalonyx, and Mylodon. Within nearly this same period (as proved by the shells at Bahia Blanca) South America possessed, as we have just seen, a mastodon, horse, hollow-horned ruminant, and the same three genera (as well as several others) of the Edentata. Hence it is evident that North and South America, in having within a late geological period these several genera in common, were much more closely related in the character of their terrestrial inhabitants than they now are. The more I reflect on this case, the more interesting it appears: I know of no other instance where we can almost mark the period and manner of the splitting up of one great region into two well-characterized zoological provinces. The geologist, who is fully impressed with the vast oscillations of level which have affected the earths crust within late periods, will not fear to speculate on the recent elevation of the Mexican platform, or, more probably, on the recent submergence of land in the West Indian Archipelago, as the cause of the present zoological separation of North and South America. The South American character of the West Indian mammals<a href="endnotes.xhtml#note-77" id="noteref-77" epub:type="noteref">77</a> seems to indicate that this archipelago was formerly united to the southern continent, and that it has subsequently been an area of subsidence.</p>
<p>When America, and especially North America, possessed its elephants, mastodons, horse, and hollow-horned ruminants, it was much more closely related in its zoological characters to the temperate parts of Europe and Asia than it now is. As the remains of these genera are found on both sides of Behrings Straits<a href="endnotes.xhtml#note-78" id="noteref-78" epub:type="noteref">78</a> and on the plains of Siberia, we are led to look to the northwestern side of North America as the former point of communication between the Old and so-called New World. And as so many species, both living and extinct, of these same genera inhabit and have inhabited the Old World, it seems most probable that the North American elephants, mastodons, horse, and hollow-horned ruminants migrated, on land since submerged near Behrings Straits, from Siberia into North America, and thence, on land since submerged in the West Indies, into South America, where for a time they mingled with the forms characteristic of that southern continent, and have since become extinct.</p> <p>When America, and especially North America, possessed its elephants, mastodons, horse, and hollow-horned ruminants, it was much more closely related in its zoological characters to the temperate parts of Europe and Asia than it now is. As the remains of these genera are found on both sides of Berings Straits<a href="endnotes.xhtml#note-78" id="noteref-78" epub:type="noteref">78</a> and on the plains of Siberia, we are led to look to the northwestern side of North America as the former point of communication between the Old and so-called New World. And as so many species, both living and extinct, of these same genera inhabit and have inhabited the Old World, it seems most probable that the North American elephants, mastodons, horse, and hollow-horned ruminants migrated, on land since submerged near Berings Straits, from Siberia into North America, and thence, on land since submerged in the West Indies, into South America, where for a time they mingled with the forms characteristic of that southern continent, and have since become extinct.</p>
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<p>While travelling through the country, I received several vivid descriptions of the effects of a late great drought; and the account of this may throw some light on the cases where vast numbers of animals of all kinds have been embedded together. The period included between the years 1827 and 1830 is called the “<i xml:lang="es">gran seco</i>,” or the great drought. During this time so little rain fell, that the vegetation, even to the thistles, failed; the brooks were dried up, and the whole country assumed the appearance of a dusty high road. This was especially the case in the northern part of the province of Buenos Aires and the southern part of <abbr>St.</abbr> Fé. Very great numbers of birds, wild animals, cattle, and horses perished from the want of food and water. A man told me that the deer<a href="endnotes.xhtml#note-79" id="noteref-79" epub:type="noteref">79</a> used to come into his courtyard to the well, which he had been obliged to dig to supply his own family with water; and that the partridges had hardly strength to fly away when pursued. The lowest estimation of the loss of cattle in the province of Buenos Aires alone, was taken at one million head. A proprietor at San Pedro had previously to these years 20,000 cattle; at the end not one remained. San Pedro is situated in the middle of the finest country; and even now abounds again with animals; yet during the latter part of the “<i xml:lang="es">gran seco</i>,” live cattle were brought in vessels for the consumption of the inhabitants. The animals roamed from their estancias, and, wandering far southward, were mingled together in such multitudes, that a government commission was sent from Buenos Aires to settle the disputes of the owners. Sir Woodbine Parish informed me of another and very curious source of dispute; the ground being so long dry, such quantities of dust were blown about, that in this open country the landmarks became obliterated, and people could not tell the limits of their estates.</p> <p>While travelling through the country, I received several vivid descriptions of the effects of a late great drought; and the account of this may throw some light on the cases where vast numbers of animals of all kinds have been embedded together. The period included between the years 1827 and 1830 is called the “<i xml:lang="es">gran seco</i>,” or the great drought. During this time so little rain fell, that the vegetation, even to the thistles, failed; the brooks were dried up, and the whole country assumed the appearance of a dusty high road. This was especially the case in the northern part of the province of Buenos Aires and the southern part of <abbr>St.</abbr> Fé. Very great numbers of birds, wild animals, cattle, and horses perished from the want of food and water. A man told me that the deer<a href="endnotes.xhtml#note-79" id="noteref-79" epub:type="noteref">79</a> used to come into his courtyard to the well, which he had been obliged to dig to supply his own family with water; and that the partridges had hardly strength to fly away when pursued. The lowest estimation of the loss of cattle in the province of Buenos Aires alone, was taken at one million head. A proprietor at San Pedro had previously to these years 20,000 cattle; at the end not one remained. San Pedro is situated in the middle of the finest country; and even now abounds again with animals; yet during the latter part of the “<i xml:lang="es">gran seco</i>,” live cattle were brought in vessels for the consumption of the inhabitants. The animals roamed from their estancias, and, wandering far southward, were mingled together in such multitudes, that a government commission was sent from Buenos Aires to settle the disputes of the owners. Sir Woodbine Parish informed me of another and very curious source of dispute; the ground being so long dry, such quantities of dust were blown about, that in this open country the landmarks became obliterated, and people could not tell the limits of their estates.</p>
<p>I was informed by an eyewitness that the cattle in herds of thousands rushed into the Parana, and being exhausted by hunger they were unable to crawl up the muddy banks, and thus were drowned. The arm of the river which runs by San Pedro was so full of putrid carcasses, that the master of a vessel told me that the smell rendered it quite impassable. Without doubt several hundred thousand animals thus perished in the river: their bodies when putrid were seen floating down the stream; and many in all probability were deposited in the estuary of the Plata. All the small rivers became highly saline, and this caused the death of vast numbers in particular spots; for when an animal drinks of such water it does not recover. Azara describes<a href="endnotes.xhtml#note-80" id="noteref-80" epub:type="noteref">80</a> the fury of the wild horses on a similar occasion, rushing into the marshes, those which arrived first being overwhelmed and crushed by those which followed. He adds that more than once he has seen the carcasses of upwards of a thousand wild horses thus destroyed. I noticed that the smaller streams in the Pampas were paved with a breccia of bones but this probably is the effect of a gradual increase, rather than of the destruction at any one period. Subsequently to the drought of 1827 to 1832, a very rainy season followed which caused great floods. Hence it is almost certain that some thousands of the skeletons were buried by the deposits of the very next year. What would be the opinion of a geologist, viewing such an enormous collection of bones, of all kinds of animals and of all ages, thus embedded in one thick earthy mass? Would he not attribute it to a flood having swept over the surface of the land, rather than to the common order of things?<a href="endnotes.xhtml#note-81" id="noteref-81" epub:type="noteref">81</a></p> <p>I was informed by an eyewitness that the cattle in herds of thousands rushed into the Parana, and being exhausted by hunger they were unable to crawl up the muddy banks, and thus were drowned. The arm of the river which runs by San Pedro was so full of putrid carcasses, that the master of a vessel told me that the smell rendered it quite impassable. Without doubt several hundred thousand animals thus perished in the river: their bodies when putrid were seen floating down the stream; and many in all probability were deposited in the estuary of the Plata. All the small rivers became highly saline, and this caused the death of vast numbers in particular spots; for when an animal drinks of such water it does not recover. Azara describes<a href="endnotes.xhtml#note-80" id="noteref-80" epub:type="noteref">80</a> the fury of the wild horses on a similar occasion, rushing into the marshes, those which arrived first being overwhelmed and crushed by those which followed. He adds that more than once he has seen the carcasses of upwards of a thousand wild horses thus destroyed. I noticed that the smaller streams in the Pampas were paved with a breccia of bones but this probably is the effect of a gradual increase, rather than of the destruction at any one period. Subsequently to the drought of 1827 to 1832, a very rainy season followed which caused great floods. Hence it is almost certain that some thousands of the skeletons were buried by the deposits of the very next year. What would be the opinion of a geologist, viewing such an enormous collection of bones, of all kinds of animals and of all ages, thus embedded in one thick earthy mass? Would he not attribute it to a flood having swept over the surface of the land, rather than to the common order of things?<a href="endnotes.xhtml#note-81" id="noteref-81" epub:type="noteref">81</a></p>

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<p>Everything in this southern continent has been effected on a grand scale: the land, from the Rio Plata to Tierra del Fuego, a distance of 1,200 miles, has been raised in mass (and in Patagonia to a height of between 300 and 400 feet), within the period of the now existing seashells. The old and weathered shells left on the surface of the upraised plain still partially retain their colours. The uprising movement has been interrupted by at least eight long periods of rest, during which the sea ate, deeply back into the land, forming at successive levels the long lines of cliffs, or escarpments, which separate the different plains as they rise like steps one behind the other. The elevatory movement, and the eating-back power of the sea during the periods of rest, have been equable over long lines of coast; for I was astonished to find that the step-like plains stand at nearly corresponding heights at far distant points. The lowest plain is 90 feet high; and the highest, which I ascended near the coast, is 950 feet; and of this, only relics are left in the form of flat gravel-capped hills. The upper plain of Santa Cruz slopes up to a height of 3,000 feet at the foot of the Cordillera. I have said that within the period of existing seashells, Patagonia has been upraised 300 to 400 feet: I may add, that within the period when icebergs transported boulders over the upper plain of Santa Cruz, the elevation has been at least 1,500 feet. Nor has Patagonia been affected only by upward movements: the extinct tertiary shells from Port <abbr>St.</abbr> Julian and Santa Cruz cannot have lived, according to Professor <abbr epub:type="z3998:given-name">E.</abbr> Forbes, in a greater depth of water than from 40 to 250 feet; but they are now covered with sea-deposited strata from 800 to 1,000 feet in thickness: hence the bed of the sea, on which these shells once lived, must have sunk downwards several hundred feet, to allow of the accumulation of the superincumbent strata. What a history of geological changes does the simply-constructed coast of Patagonia reveal!</p> <p>Everything in this southern continent has been effected on a grand scale: the land, from the Rio Plata to Tierra del Fuego, a distance of 1,200 miles, has been raised in mass (and in Patagonia to a height of between 300 and 400 feet), within the period of the now existing seashells. The old and weathered shells left on the surface of the upraised plain still partially retain their colours. The uprising movement has been interrupted by at least eight long periods of rest, during which the sea ate, deeply back into the land, forming at successive levels the long lines of cliffs, or escarpments, which separate the different plains as they rise like steps one behind the other. The elevatory movement, and the eating-back power of the sea during the periods of rest, have been equable over long lines of coast; for I was astonished to find that the step-like plains stand at nearly corresponding heights at far distant points. The lowest plain is 90 feet high; and the highest, which I ascended near the coast, is 950 feet; and of this, only relics are left in the form of flat gravel-capped hills. The upper plain of Santa Cruz slopes up to a height of 3,000 feet at the foot of the Cordillera. I have said that within the period of existing seashells, Patagonia has been upraised 300 to 400 feet: I may add, that within the period when icebergs transported boulders over the upper plain of Santa Cruz, the elevation has been at least 1,500 feet. Nor has Patagonia been affected only by upward movements: the extinct tertiary shells from Port <abbr>St.</abbr> Julian and Santa Cruz cannot have lived, according to Professor <abbr epub:type="z3998:given-name">E.</abbr> Forbes, in a greater depth of water than from 40 to 250 feet; but they are now covered with sea-deposited strata from 800 to 1,000 feet in thickness: hence the bed of the sea, on which these shells once lived, must have sunk downwards several hundred feet, to allow of the accumulation of the superincumbent strata. What a history of geological changes does the simply-constructed coast of Patagonia reveal!</p>
<p>At Port <abbr>St.</abbr> Julian,<a href="endnotes.xhtml#note-93" id="noteref-93" epub:type="noteref">93</a> in some red mud capping the gravel on the 90-feet plain, I found half the skeleton of the <i epub:type="z3998:taxonomy">Macrauchenia Patachonica</i>, a remarkable quadruped, full as large as a camel. It belongs to the same division of the Pachydermata with the rhinoceros, tapir, and palaeotherium; but in the structure of the bones of its long neck it shows a clear relation to the camel, or rather to the guanaco and llama. From recent seashells being found on two of the higher step-formed plains, which must have been modelled and upraised before the mud was deposited in which the macrauchenia was entombed, it is certain that this curious quadruped lived long after the sea was inhabited by its present shells. I was at first much surprised how a large quadruped could so lately have subsisted, in <abbr>lat.</abbr> 49° 15, on these wretched gravel plains, with their stunted vegetation; but the relationship of the Macrauchenia to the Guanaco, now an inhabitant of the most sterile parts, partly explains this difficulty.</p> <p>At Port <abbr>St.</abbr> Julian,<a href="endnotes.xhtml#note-93" id="noteref-93" epub:type="noteref">93</a> in some red mud capping the gravel on the 90-feet plain, I found half the skeleton of the <i epub:type="z3998:taxonomy">Macrauchenia Patachonica</i>, a remarkable quadruped, full as large as a camel. It belongs to the same division of the Pachydermata with the rhinoceros, tapir, and palaeotherium; but in the structure of the bones of its long neck it shows a clear relation to the camel, or rather to the guanaco and llama. From recent seashells being found on two of the higher step-formed plains, which must have been modelled and upraised before the mud was deposited in which the macrauchenia was entombed, it is certain that this curious quadruped lived long after the sea was inhabited by its present shells. I was at first much surprised how a large quadruped could so lately have subsisted, in <abbr>lat.</abbr> 49° 15, on these wretched gravel plains, with their stunted vegetation; but the relationship of the Macrauchenia to the Guanaco, now an inhabitant of the most sterile parts, partly explains this difficulty.</p>
<p>The relationship, though distant, between the Macrauchenia and the Guanaco, between the Toxodon and the Capybara—the closer relationship between the many extinct Edentata and the living sloths, anteaters, and armadillos, now so eminently characteristic of South American zoology—and the still closer relationship between the fossil and living species of Ctenomys and Hydrochaerus, are most interesting facts. This relationship is shown wonderfully—as wonderfully as between the fossil and extinct marsupial animals of Australia—by the great collection lately brought to Europe from the caves of Brazil by <abbr epub:type="z3998:name-title">MM.</abbr> Lund and Clausen. In this collection there are extinct species of all the thirty-two genera, excepting four, of the terrestrial quadrupeds now inhabiting the provinces in which the caves occur; and the extinct species are much more numerous than those now living: there are fossil anteaters, armadillos, tapirs, peccaries, guanacos, opossums, and numerous South American gnawers and monkeys, and other animals. This wonderful relationship in the same continent between the dead and the living, will, I do not doubt, hereafter throw more light on the appearance of organic beings on our earth, and their disappearance from it, than any other class of facts.</p> <p>The relationship, though distant, between the Macrauchenia and the Guanaco, between the Toxodon and the Capybara—the closer relationship between the many extinct Edentata and the living sloths, anteaters, and armadillos, now so eminently characteristic of South American zoology—and the still closer relationship between the fossil and living species of Ctenomys and Hydrochaerus, are most interesting facts. This relationship is shown wonderfully—as wonderfully as between the fossil and extinct marsupial animals of Australia—by the great collection lately brought to Europe from the caves of Brazil by <abbr epub:type="z3998:name-title">MM.</abbr> Lund and Clausen. In this collection there are extinct species of all the thirty-two genera, excepting four, of the terrestrial quadrupeds now inhabiting the provinces in which the caves occur; and the extinct species are much more numerous than those now living: there are fossil anteaters, armadillos, tapirs, peccaries, guanacos, opossums, and numerous South American gnawers and monkeys, and other animals. This wonderful relationship in the same continent between the dead and the living, will, I do not doubt, hereafter throw more light on the appearance of organic beings on our earth, and their disappearance from it, than any other class of facts.</p>
<p>It is impossible to reflect on the changed state of the American continent without the deepest astonishment. Formerly it must have swarmed with great monsters: now we find mere pygmies, compared with the antecedent, allied races. If Buffon had known of the gigantic sloth and armadillo-like animals, and of the lost Pachydermata, he might have said with a greater semblance of truth that the creative force in America had lost its power, rather than that it had never possessed great vigour. The greater number, if not all, of these extinct quadrupeds lived at a late period, and were the contemporaries of most of the existing seashells. Since they lived, no very great change in the form of the land can have taken place. What, then, has exterminated so many species and whole genera? The mind at first is irresistibly hurried into the belief of some great catastrophe; but thus to destroy animals, both large and small, in Southern Patagonia, in Brazil, on the Cordillera of Peru, in North America up to Behrings Straits, we must shake the entire framework of the globe. An examination, moreover, of the geology of La Plata and Patagonia, leads to the belief that all the features of the land result from slow and gradual changes. It appears from the character of the fossils in Europe, Asia, Australia, and in North and South America, that those conditions which favour the life of the <em>larger</em> quadrupeds were lately coextensive with the world: what those conditions were, no one has yet even conjectured. It could hardly have been a change of temperature, which at about the same time destroyed the inhabitants of tropical, temperate, and arctic latitudes on both sides of the globe. In North America we positively know from <abbr epub:type="z3998:name-title">Mr.</abbr> Lyell, that the large quadrupeds lived subsequently to that period, when boulders were brought into latitudes at which icebergs now never arrive: from conclusive but indirect reasons we may feel sure, that in the southern hemisphere the Macrauchenia, also, lived long subsequently to the ice-transporting boulder-period. Did man, after his first inroad into South America, destroy, as has been suggested, the unwieldy Megatherium and the other Edentata? We must at least look to some other cause for the destruction of the little tucutuco at Bahia Blanca, and of the many fossil mice and other small quadrupeds in Brazil. No one will imagine that a drought, even far severer than those which cause such losses in the provinces of La Plata, could destroy every individual of every species from Southern Patagonia to Behrings Straits. What shall we say of the extinction of the horse? Did those plains fail of pasture, which have since been overrun by thousands and hundreds of thousands of the descendants of the stock introduced by the Spaniards? Have the subsequently introduced species consumed the food of the great antecedent races? Can we believe that the Capybara has taken the food of the Toxodon, the Guanaco of the Macrauchenia, the existing small Edentata of their numerous gigantic prototypes? Certainly, no fact in the long history of the world is so startling as the wide and repeated exterminations of its inhabitants.</p> <p>It is impossible to reflect on the changed state of the American continent without the deepest astonishment. Formerly it must have swarmed with great monsters: now we find mere pygmies, compared with the antecedent, allied races. If Buffon had known of the gigantic sloth and armadillo-like animals, and of the lost Pachydermata, he might have said with a greater semblance of truth that the creative force in America had lost its power, rather than that it had never possessed great vigour. The greater number, if not all, of these extinct quadrupeds lived at a late period, and were the contemporaries of most of the existing seashells. Since they lived, no very great change in the form of the land can have taken place. What, then, has exterminated so many species and whole genera? The mind at first is irresistibly hurried into the belief of some great catastrophe; but thus to destroy animals, both large and small, in Southern Patagonia, in Brazil, on the Cordillera of Peru, in North America up to Berings Straits, we must shake the entire framework of the globe. An examination, moreover, of the geology of La Plata and Patagonia, leads to the belief that all the features of the land result from slow and gradual changes. It appears from the character of the fossils in Europe, Asia, Australia, and in North and South America, that those conditions which favour the life of the <em>larger</em> quadrupeds were lately coextensive with the world: what those conditions were, no one has yet even conjectured. It could hardly have been a change of temperature, which at about the same time destroyed the inhabitants of tropical, temperate, and arctic latitudes on both sides of the globe. In North America we positively know from <abbr epub:type="z3998:name-title">Mr.</abbr> Lyell, that the large quadrupeds lived subsequently to that period, when boulders were brought into latitudes at which icebergs now never arrive: from conclusive but indirect reasons we may feel sure, that in the southern hemisphere the Macrauchenia, also, lived long subsequently to the ice-transporting boulder-period. Did man, after his first inroad into South America, destroy, as has been suggested, the unwieldy Megatherium and the other Edentata? We must at least look to some other cause for the destruction of the little tucutuco at Bahia Blanca, and of the many fossil mice and other small quadrupeds in Brazil. No one will imagine that a drought, even far severer than those which cause such losses in the provinces of La Plata, could destroy every individual of every species from Southern Patagonia to Berings Straits. What shall we say of the extinction of the horse? Did those plains fail of pasture, which have since been overrun by thousands and hundreds of thousands of the descendants of the stock introduced by the Spaniards? Have the subsequently introduced species consumed the food of the great antecedent races? Can we believe that the Capybara has taken the food of the Toxodon, the Guanaco of the Macrauchenia, the existing small Edentata of their numerous gigantic prototypes? Certainly, no fact in the long history of the world is so startling as the wide and repeated exterminations of its inhabitants.</p>
<p>Nevertheless, if we consider the subject under another point of view, it will appear less perplexing. We do not steadily bear in mind, how profoundly ignorant we are of the conditions of existence of every animal; nor do we always remember, that some check is constantly preventing the too rapid increase of every organized being left in a state of nature. The supply of food, on an average, remains constant, yet the tendency in every animal to increase by propagation is geometrical; and its surprising effects have nowhere been more astonishingly shown, than in the case of the European animals run wild during the last few centuries in America. Every animal in a state of nature regularly breeds; yet in a species long established, any <em>great</em> increase in numbers is obviously impossible, and must be checked by some means. We are, nevertheless, seldom able with certainty to tell in any given species, at what period of life, or at what period of the year, or whether only at long intervals, the check falls; or, again, what is the precise nature of the check. Hence probably it is, that we feel so little surprise at one, of two species closely allied in habits, being rare and the other abundant in the same district; or, again, that one should be abundant in one district, and another, filling the same place in the economy of nature, should be abundant in a neighbouring district, differing very little in its conditions. If asked how this is, one immediately replies that it is determined by some slight difference, in climate, food, or the number of enemies: yet how rarely, if ever, we can point out the precise cause and manner of action of the check! We are therefore, driven to the conclusion, that causes generally quite inappreciable by us, determine whether a given species shall be abundant or scanty in numbers.</p> <p>Nevertheless, if we consider the subject under another point of view, it will appear less perplexing. We do not steadily bear in mind, how profoundly ignorant we are of the conditions of existence of every animal; nor do we always remember, that some check is constantly preventing the too rapid increase of every organized being left in a state of nature. The supply of food, on an average, remains constant, yet the tendency in every animal to increase by propagation is geometrical; and its surprising effects have nowhere been more astonishingly shown, than in the case of the European animals run wild during the last few centuries in America. Every animal in a state of nature regularly breeds; yet in a species long established, any <em>great</em> increase in numbers is obviously impossible, and must be checked by some means. We are, nevertheless, seldom able with certainty to tell in any given species, at what period of life, or at what period of the year, or whether only at long intervals, the check falls; or, again, what is the precise nature of the check. Hence probably it is, that we feel so little surprise at one, of two species closely allied in habits, being rare and the other abundant in the same district; or, again, that one should be abundant in one district, and another, filling the same place in the economy of nature, should be abundant in a neighbouring district, differing very little in its conditions. If asked how this is, one immediately replies that it is determined by some slight difference, in climate, food, or the number of enemies: yet how rarely, if ever, we can point out the precise cause and manner of action of the check! We are therefore, driven to the conclusion, that causes generally quite inappreciable by us, determine whether a given species shall be abundant or scanty in numbers.</p>
<p>In the cases where we can trace the extinction of a species through man, either wholly or in one limited district, we know that it becomes rarer and rarer, and is then lost: it would be difficult to point out any just distinction<a href="endnotes.xhtml#note-94" id="noteref-94" epub:type="noteref">94</a> between a species destroyed by man or by the increase of its natural enemies. The evidence of rarity preceding extinction, is more striking in the successive tertiary strata, as remarked by several able observers; it has often been found that a shell very common in a tertiary stratum is now most rare, and has even long been thought extinct. If then, as appears probable, species first become rare and then extinct—if the too rapid increase of every species, even the most favoured, is steadily checked, as we must admit, though how and when it is hard to say—and if we see, without the smallest surprise, though unable to assign the precise reason, one species abundant and another closely allied species rare in the same district—why should we feel such great astonishment at the rarity being carried one step further to extinction? An action going on, on every side of us, and yet barely appreciable, might surely be carried a little further, without exciting our observation. Who would feel any great surprise at hearing that the Magalonyx was formerly rare compared with the Megatherium, or that one of the fossil monkeys was few in number compared with one of the now living monkeys? and yet in this comparative rarity, we should have the plainest evidence of less favourable conditions for their existence. To admit that species generally become rare before they become extinct—to feel no surprise at the comparative rarity of one species with another, and yet to call in some extraordinary agent and to marvel greatly when a species ceases to exist, appears to me much the same as to admit that sickness in the individual is the prelude to death—to feel no surprise at sickness—but when the sick man dies to wonder, and to believe that he died through violence.</p> <p>In the cases where we can trace the extinction of a species through man, either wholly or in one limited district, we know that it becomes rarer and rarer, and is then lost: it would be difficult to point out any just distinction<a href="endnotes.xhtml#note-94" id="noteref-94" epub:type="noteref">94</a> between a species destroyed by man or by the increase of its natural enemies. The evidence of rarity preceding extinction, is more striking in the successive tertiary strata, as remarked by several able observers; it has often been found that a shell very common in a tertiary stratum is now most rare, and has even long been thought extinct. If then, as appears probable, species first become rare and then extinct—if the too rapid increase of every species, even the most favoured, is steadily checked, as we must admit, though how and when it is hard to say—and if we see, without the smallest surprise, though unable to assign the precise reason, one species abundant and another closely allied species rare in the same district—why should we feel such great astonishment at the rarity being carried one step further to extinction? An action going on, on every side of us, and yet barely appreciable, might surely be carried a little further, without exciting our observation. Who would feel any great surprise at hearing that the Magalonyx was formerly rare compared with the Megatherium, or that one of the fossil monkeys was few in number compared with one of the now living monkeys? and yet in this comparative rarity, we should have the plainest evidence of less favourable conditions for their existence. To admit that species generally become rare before they become extinct—to feel no surprise at the comparative rarity of one species with another, and yet to call in some extraordinary agent and to marvel greatly when a species ceases to exist, appears to me much the same as to admit that sickness in the individual is the prelude to death—to feel no surprise at sickness—but when the sick man dies to wonder, and to believe that he died through violence.</p>
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